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Defects in many developmental pathways can be classified as loss-of-function or gain-of-function. Loss-of-function mutations inactivating the gene are dominant and most often cause lethality and embryonic death. Gain-of-function mutations are usually recessive and can cause a wide range of phenotypes. The Drosophila singed locus encodes a protein necessary for multiple embryonic developmental processes. In this study, we analyzed mutations that are associated with the singed locus for possible gain-of-function phenotypes. Several new alleles of singed that cause lethality or adult-onset defects were identified and the phenotypes were mapped to a genomic region on chromosome 2. Two new alleles of singed also had profound effects on localization of the homeotic gene Distalless. These results demonstrate that singed encodes a protein that has different functions during different stages of embryonic development.Burkhard Krüger

Burkhard Krüger (born 14 June 1967) is a retired German football player. He is now a goalkeeping coach for his former team 1. FC Saarbrücken.

Club career
During his playing career he played for 1. FC Saarbrücken, AS Saint-Étienne, Karlsruher SC, KFC Uerdingen 05, Rot Weiss Ahlen, SSV Reutlingen, Alemannia Aachen and Carl Zeiss Jena.

He made his debut on 7 May 1985 in a DFB-Pokal match against SC Freiburg. His last game was on 1 June 1998 in the 2. Bundesliga match for Rot Weiss Ahlen against V


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